1984, MS, Biophysics, Shevchenko Kiev State University, Kiev, Ukraine  1989, PhD, Oncology, Kavetsky Institute of Oncology Problems, Kiev, Academy of Science of the USSR  1989-1992 Postdoc, Cancer Biology,  Kavetsky Institute of Oncology Problems, Kiev, Ukraine  1992-1996, Postdoc, University of Arizona Cancer Center, Tucson, Arizona, USA 1996-1999, Assistant Scientific Investigator, University of Arizona cancer center, Tucson, AZ  1999-2003, Assistant Professor (research track) Department of Radiation Oncology, University of Arizona Cancer Center, Tucson, Arizona  2003- 2008 Assistant Professor (research track),  Department of Cell Biology and Anatomy, College of Medicine, University of Arizona, Tucson, Arizona  2009-present, Associate Professor (Research  Scholar Track, career Track) , Department of Cellular and Molecular Medicine, College of Medicine, University of Arizona, Tucson, Arizona